For serious wounding by an infiltrator, Council positions are lost instantly, except the Ruler position in a Monarch, Theocracy, or Tyranny.
All positions can be lost if the character remains wounded for long enough, from any kind of wound.
We know that there is a certain amount of inconsistency in how wounding is treated. We intend to clear it up, but due to the way the code in these situations works, it's not as simple as it sounds.